Sea Snail
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Sea snail is a common name for slow-moving marine gastropod
molluscs Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is estim ...
, usually with visible external shells, such as
whelk Whelk (also known as scungilli) is a common name applied to various kinds of sea snail. Although a number of whelks are relatively large and are in the family Buccinidae (the true whelks), the word ''whelk'' is also applied to some other marin ...
or abalone. They share the taxonomic class Gastropoda with
slugs Slug, or land slug, is a common name for any apparently shell-less terrestrial gastropod mollusc. The word ''slug'' is also often used as part of the common name of any gastropod mollusc that has no shell, a very reduced shell, or only a sma ...
, which are distinguished from
snails A snail is, in loose terms, a shelled gastropod. The name is most often applied to land snails, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class Gastro ...
primarily by the absence of a visible
shell Shell may refer to: Architecture and design * Shell (structure), a thin structure ** Concrete shell, a thin shell of concrete, usually with no interior columns or exterior buttresses ** Thin-shell structure Science Biology * Seashell, a hard o ...
.


Definition

Determining whether some gastropods should be called sea snails is not always easy. Some species that live in
brackish water Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
(such as certain neritids) can be listed as either freshwater snails or marine snails, and some species that live at or just above the
high tide Tides are the rise and fall of sea levels caused by the combined effects of the gravitational forces exerted by the Moon (and to a much lesser extent, the Sun) and are also caused by the Earth and Moon orbiting one another. Tide tables c ...
level (for example species in the genus '' Truncatella'') are sometimes considered to be sea snails and sometimes listed as
land snails A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial gastropod mollusks that have shells (those without shells are known as ...
.


Anatomy

Sea snails are a very large group of animals and a very diverse one. Most snails that live in
salt water Saline water (more commonly known as salt water) is water that contains a high concentration of dissolved salts (mainly sodium chloride). On the United States Geological Survey (USGS) salinity scale, saline water is saltier than brackish wat ...
respire using a
gill A gill () is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they are ...
or gills; a few species, though, have a lung, are
intertidal The intertidal zone, also known as the foreshore, is the area above water level at low tide and underwater at high tide (in other words, the area within the tidal range). This area can include several types of habitats with various species ...
, and are active only at low tide when they can move around in the air. These air-breathing species include false limpets in the family
Siphonariidae Siphonariidae, also known as false limpets, are a taxonomic family of small to medium-sized air-breathing sea snails, marine and brackish water pulmonate gastropod molluscs. __TOC__ Shell description Empty siphonariid limpet shells can be dis ...
and another group of false limpets in the family
Trimusculidae ''Trimusculus'' is a genus of medium-sized air-breathing sea snails or false limpets, marine pulmonate gastropod molluscs in the family Trimusculidae. ''Trimusculus'' is the only genus in the family Trimusculidae. Trimusculidae, the butt ...
. Many, but not all, sea snails have an operculum.


Shell

The shells of most species of sea snails are spirally coiled. Some, though, have conical shells, and these are often referred to by the common name of limpets. In one unusual family (
Juliidae Juliidae, common name the bivalved gastropods, is a family of minute sea snails, marine gastropod mollusks or micromollusks in the superfamily Oxynooidea, an opisthobranch group. MolluscaBase eds. (2021). MolluscaBase. Juliidae E. A. Smith ...
), the shell of the snail has become two hinged plates closely resembling those of a
bivalve Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bival ...
; this family is sometimes called the "bivalved gastropods". Their shells are found in a variety of shapes and sizes, but are normally very small. Those of living species of sea snails range in size from '' Syrinx aruanus'', the largest living shelled gastropod species at , to minute species whose shells are less than 1 mm at adult size. Because the shells of sea snails are strong and durable in many cases, as a group they are well represented in the
fossil record A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
. The shells of snails are complex and grow at different speeds. The speed of growth is affected by a few variables such as the temperature of the water, depth of the water, food present for the snail, as well as isotopic oxygen levels. By looking at the composition of
aragonite Aragonite is a carbonate mineral, one of the three most common naturally occurring crystal forms of calcium carbonate, (the other forms being the minerals calcite and vaterite). It is formed by biological and physical processes, including pre ...
in the growth layers of mollusks you can predict the size the mollusk shell can reach.


Taxonomy


2005 taxonomy

The following
cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...
is an overview of the main clades of living gastropods based on the taxonomy of Bouchet & Rocroi (2005), with taxa that contain saltwater or brackish water species marked in
boldface In typography, emphasis is the strengthening of words in a text with a font in a different style from the rest of the text, to highlight them. It is the equivalent of prosody stress in speech. Methods and use The most common methods in W ...
(some of the highlighted taxa consist entirely of marine species, but some of them also contain freshwater or land species.) * Clade
Patellogastropoda The Patellogastropoda, common name true limpets and historically called the Docoglossa, are members of a major phylogenetic group of marine gastropods, treated by experts either as a clade or as a taxonomic order. The clade Patellogastropoda is d ...
* Clade
Vetigastropoda Vetigastropoda is a major taxonomic group of sea snails, marine gastropod mollusks that form a very ancient lineage. Taxonomically the Vetigastropoda are sometimes treated as an order, although they are treated as an unranked clade in Bouchet ...
* Clade Cocculiniformia * Clade
Neritimorpha Neritimorpha is a taxonomic grouping, an unranked major clade of snails, gastropod mollusks. This grouping includes land snails, sea snails, slugs, some deepwater limpets, and also freshwater snails. Neritimorpha contains around 2,000 extant s ...
** Clade
Cycloneritimorpha Cycloneritida (nerites and false-limpets) is an order of land snails, freshwater snails, and sea snails.MolluscaBase eds. (2020). MolluscaBase. Cycloneritida. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/ap ...
* Clade Caenogastropoda ** Informal group Architaenioglossa ** Clade
Sorbeoconcha Sorbeoconcha is a taxonomic clade of snails, i.e. gastropods, mainly marine species with gills and opercula, within the clade Caenogastropoda. The taxon Sorbeoconcha was named by Winston Ponder and David R. Lindberg in 1997. Taxonomy 1 ...
** Clade Hypsogastropoda *** Clade
Littorinimorpha Littorinimorpha is a large order of snails, gastropods, consisting primarily of sea snails (marine species), but also including some freshwater snails ( aquatic species) and land snails ( terrestrial species).Bouchet P. & Rocroi J.-P. (Ed.); Frý ...
*** Informal group
Ptenoglossa The Ptenoglossa is an informal taxonomic group of sea snails. This group was considered paraphyletic or polyphyletic by Ponder and Lindberg (1997) in their classification of gastropod molluscs. The database WoRMS considers this suborder as b ...
*** Clade Neogastropoda * Clade
Heterobranchia Heterobranchia, the ''heterobranchs'' (meaning "different-gilled snails"), is a taxonomic clade of snails and slugs, which includes marine, aquatic and terrestrial gastropod mollusks. Heterobranchia is one of the main clades of gastropods. Cur ...
** Informal group
Lower Heterobranchia Lower Heterobranchia, also known as the Allogastropoda, is a group of rather specialized, highly evolved sea slugs and sea snails, ( marine gastropod mollusks) within the subclass Heterobranchia. WoRMS (2020). "Lower Heterobranchia". Accessed at ...
** Informal group Opisthobranchia *** Clade
Cephalaspidea The order Cephalaspidea, also known as the headshield slugs and bubble snails, is a major taxon of sea slugs and bubble snails, marine gastropod mollusks within the larger clade Euopisthobranchia.Jörger K. M., Stöger I., Kano Y., Fukuda H., ...
*** Clade
Thecosomata Sea butterflies, scientific name Thecosomata (thecosomes, "case / shell-body"), are a taxonomic suborder of small pelagic swimming sea snails. They are holoplanktonic opisthobranch gastropod mollusks. Most Thecosomata have some form of calcifi ...
*** Clade
Gymnosomata Sea angels (clade Gymnosomata) are a large group of small free-swimming sea slugs, not to be confused with Cnidarians (Jellyfish and other similar creatures), classified into six different families. They are pelagic opisthobranchs in the clade Gy ...
*** Clade
Aplysiomorpha The clade Anaspidea, commonly known as sea hares (''Aplysia'' species and related genera), are medium-sized to very large opisthobranch gastropod molluscs with a soft internal shell made of protein. These are marine gastropod molluscs in the s ...
*** Group
Acochlidiacea Acochlidiacea, common name acochlidians, are a taxonomic clade of very unusual sea snails and sea and freshwater slugs, aquatic gastropod mollusks within the large clade Heterobranchia. Acochlidia is a variant spelling. Description These a ...
*** Clade
Sacoglossa Sacoglossa, commonly known as the sacoglossans or the "solar-powered sea slugs", are a superorder of small sea slugs and sea snails, marine gastropod mollusks that belong to the clade Heterobranchia. Sacoglossans live by ingesting the cellular co ...
*** Group Cylindrobullida *** Clade
Umbraculida Umbraculoidea is a superfamily of unusual false limpets with a thin soft patelliform shell, marine gastropod molluscs in the clade Umbraculida, within the clade Euopisthobranchia. There are two families in this superfamily, which is listed as ...
*** Clade
Nudipleura Nudipleura are a clade of sea snails and sea slugs, marine gastropod mollusks within the large clade Heterobranchia. Taxonomy 2005 taxonomy In the taxonomy of Bouchet & Rocroi (2005), the clade Nudipleura is arranged as follows: *Clade Nud ...
**** Clade
Pleurobranchomorpha The Pleurobranchidae are a taxonomic family of sea slugs, marine gastropod molluscs in the clade Pleurobranchomorpha. Characteristics Species in the family Pleurobranchidae have a prominent mantle and an internal shell that becomes reduced ...
**** Clade Nudibranchia ***** Clade
Euctenidiacea The Euctenidiacea, common name dorid nudibranchs, are a taxonomic suborder of sea snails or slugs, marine gastropod molluscs in the order Nudibranchia. Bouchet & Rocroi (2005) rejected the name Anthobranchia on the grounds that it also included ...
***** Clade
Dexiarchia The Dexiarchia are a suborder of sea slugs, shell-less marine gastropod molluscs in the order Nudibranchia. This classification is based on the study by Schrödl ''et al.,'' published in 2001, who recognized within this clade two clades Pseudoeu ...
****** Clade
Pseudoeuctenidiacea Doridoxidae, commonly known as doridoxid nudibranchs, are a small, enigmatic taxonomic family of shell-less sea slugs, marine gastropod molluscs in the clade Cladobranchia within the clade Nudibranchia. Doridoxidae is currently the only fa ...
****** Clade
Cladobranchia The Cladobranchia are a taxonomic clade of nudibranchs, sea slugs, marine gastropod molluscs in the clade Dexiarchia. Taxonomy clade Metarminida * Superfamily Metarminoidea This name is not available as a Superfamily name as it is not based o ...
******* Clade
Euarminida Arminoidea is a superfamily of small sea slugs, marine gastropod molluscs in the clade Nudibranchia. Arminoidea is the only superfamily in the clade Euarminida. Families A study, published in 2000, has demonstrated that Arminoidea is parap ...
******* Clade
Dendronotida Dendronotoidea is a taxonomic superfamily of small colorful sea slugs or nudibranchs, aeolid nudibranchs. MolluscaBase eds. (2022). MolluscaBase. Dendronotoidea Allman, 1845. Accessed through: World Register of Marine Species at: http://www.m ...
******* Clade
Aeolidida The Aeolidida is a taxonomic clade of sea slugs, specifically aeolid nudibranchs, marine gastropod molluscs in the clade Cladobranchia. They are distinguished from other nudibranchs by their possession of cerata containing cnidosacs. Taxo ...
** Informal group Pulmonata *** Informal group Basommatophora *** Clade
Eupulmonata Eupulmonata is a taxonomic clade of air-breathing snails. The great majority of this group are land snails and slugs, but some are marine and some are saltmarsh snails that can tolerate salty conditions. Linnean taxonomy *Suborder Eupulmonat ...
**** Clade Systellommatophora **** Clade Stylommatophora ***** Clade
Elasmognatha Elasmognatha is a taxonomic grouping, a clade, of air-breathing land snails and slugs, terrestrial pulmonate gastropod molluscs. A number of species in this taxonomic group are endangered. Taxonomy In the taxonomy of the Gastropoda by Bou ...
***** Clade
Orthurethra Orthurethra is a clade of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the clade Stylommatophora. In the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005, Orthurethra is treated as an informal group and subcla ...
***** Informal group
Sigmurethra Sigmurethra is a taxonomic category of air-breathing land snails and slugs, terrestrial pulmonate gastropod molluscs. This is an informal group which includes most land snails and slugs. The two strong synapomorphies of Sigmurethra are a ...


Uses


By humans

A number of species of sea snails are harvested in aquaculture and used by humans for food, including abalone, conch, limpets,
whelk Whelk (also known as scungilli) is a common name applied to various kinds of sea snail. Although a number of whelks are relatively large and are in the family Buccinidae (the true whelks), the word ''whelk'' is also applied to some other marin ...
s (such as the North American '' Busycon'' species and the North Atlantic ''
Buccinum undatum ''Buccinum undatum'', the common whelk or the waved buccinum, is a large, edible marine gastropod in the family Buccinidae, the "true whelks".Fraussen, K.; Gofas, S. (2014). Buccinum undatum Linnaeus, 1758. Accessed through: World Register of M ...
'') and periwinkles including ''
Littorina littorea The common periwinkle or winkle (''Littorina littorea'') is a species of small edible whelk or sea snail, a marine gastropod mollusc that has gills and an operculum, and is classified within the family Littorinidae, the periwinkles.Reid, David ...
''. The shells of sea snails are often found washed up on
beach A beach is a landform alongside a body of water which consists of loose particles. The particles composing a beach are typically made from rock, such as sand, gravel, shingle, pebbles, etc., or biological sources, such as mollusc sh ...
es. Because many are attractive and durable, they have been used to make necklaces and other
jewelry Jewellery ( UK) or jewelry ( U.S.) consists of decorative items worn for personal adornment, such as brooches, rings, necklaces, earrings, pendants, bracelets, and cufflinks. Jewellery may be attached to the body or the clothes. From a w ...
since prehistoric times. The shells of a few species of large sea snails within the
Vetigastropoda Vetigastropoda is a major taxonomic group of sea snails, marine gastropod mollusks that form a very ancient lineage. Taxonomically the Vetigastropoda are sometimes treated as an order, although they are treated as an unranked clade in Bouchet ...
have a thick layer of nacre and have been used as a source of
mother of pearl Nacre ( , ), also known as mother of pearl, is an organicinorganic composite material produced by some molluscs as an inner shell layer; it is also the material of which pearls are composed. It is strong, resilient, and iridescent. Nacre is f ...
. Historically, the
button A button is a fastener that joins two pieces of fabric together by slipping through a loop or by sliding through a buttonhole. In modern clothing and fashion design, buttons are commonly made of plastic but also may be made of metal, wood, o ...
industry relied on these species for a number of years.


By non-human animals

The shells of sea snails are used for protection by many kinds of
hermit crab Hermit crabs are anomuran decapod crustaceans of the superfamily Paguroidea that have adapted to occupy empty scavenged mollusc shells to protect their fragile exoskeletons. There are over 800 species of hermit crab, most of which possess an as ...
s. A hermit crab carries the shell by grasping the central
columella Lucius Junius Moderatus Columella (; Arabic: , 4 – ) was a prominent writer on agriculture in the Roman Empire. His ' in twelve volumes has been completely preserved and forms an important source on Roman agriculture, together with the wo ...
of the shell using claspers on the tip of its abdomen.


See also

* Freshwater snail *
Terrestrial molluscs Terrestrial molluscs or land molluscs (mollusks) are an ecological group that includes all molluscs that live on land in contrast to freshwater and marine molluscs. They probably first occurred in the Carboniferous, arising from freshwater on ...
* Land snail *
Sea slug Sea slug is a common name for some marine invertebrates with varying levels of resemblance to terrestrial slugs. Most creatures known as sea slugs are gastropods, i.e. they are sea snails (marine gastropod mollusks) that over evolutionary time ...
* Slug


References

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